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Sandalwood Plantations – Points to Ponder


Affiliations
1 Division of Farm Forestry, College of Agriculture, Bheemarayanagudi, Yadgir 585 287, India
2 Directorate of Extension, University of Agricultural Sciences, Raichur 584 104, India
 

East Indian sandalwood (Santalum album L.) is one of the precious woods known for sweet fragrant aroma and commercial value in both national and international markets. Besides, its wide adaptability to varied climate, hosts and edaphic conditions have attracted farmers and corporates for commercial venture. Sandalwood being a hemi-ischolar_main parasite, the successful establishment of sandalwood plantation depends on the understanding of parasitism ecology, especially the relations between host and parasite, their ratio and other silviculture techniques. Hence, the present article aims to understand these issues for commercial upscaling. Sandalwood relies largely on host plants for mineral nutrients and water through haustorial connections which act as a physiological and physical bridge between the parasite and the host. It parasitizes over a vast array of plants from grasses to trees, whereas leguminous associations are superior. Deepischolar_mained and slow-growing perennial hosts help in sustained growth. For better yield of heartwood and oil sandalwood should be grown over 15 years, whereas optimum rotation age would be 25–30 years. Sandalwood can be planted either through direct dibbling of seeds near the hosts in the same pit or adjacent to the hosts. Better establishment occurs when sandalwood is grown with potted host followed by secondary hosts in the field. Association of sandalwood and host would be strong when the distance is around 2.5–3 m with host to parasitic ratio of 2 : 1 or 3 : 1. However, questions on parasitism ecology still remain.

Keywords

Ecology, Haustoria, Hosts, Parasite, Sandalwood.
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  • Nageswara Rao, M., Ganeshaiah, K. N. and Uma Shaanker, R., Assessing threats and mapping sandal (Santalum album L.) resources in peninsular India: identification of genetic-hotspot for in situ conservation. Conserv. Genet., 2007, 8, 925–935.

  • Meera, C., Nageswara Rao, M., Ganeshaiah, K. N., Uma Shaanker, R. and Swaminath, M. H., Conservation of sandal genetic resources in India: I Extraction patterns and threats to sandal resources in Karnataka. My For., 2000, 36,125–132.

  • Sreenivasan, V. V., Shivaramakrishnana, V. R., Rangaswamy, C. R., Anathapadmanabha, H. S. and Shankaranarayana, K. H., Sandal, ICFRE, Dehra Dun, 1992, p. 233.

  • Suma, T. B. and Balasundaran, M., Isozyme variation in five provenances of Santalum album in India. A. J. Bot., 2003, 51, 243–249.

  • Gairola, S., Aggarwal, P. S. and Ravikumar, G. S., Status of production and marketing of sandalwood (Santalum album L.). In Proceedings of the National Seminar on Conservation, Improvement, Cultivation and Management of Sandal (eds Gairola, S. et al.), 12–13 December 2007, pp. 1–8.

  • Gillieson, D., Page, T. and Silverman, J., An Inventory of Wild Sandalwood Stocks in Vanuatu, Australian Centre for International Agricultural Research, Canberra. 2008, p. 118.

  • Subasinghe, S. M. C. U. P., Sandalwood research: a global perspective. J. Trop. For. Environ., 2013, 3, 1–8.

  • Baldovini, N., Delasalle, C. and Joulain, D., Phytochemistry of the heartwood from fragrant santalum species: a review. Flavour Frag. J., 2011, 26, 7–26.

  • Surendran, C., Parthiban, K. L., Bhuvaneswaran, C. and Murgesh, M., Silvicultural strategies for augmentation of sandal regeneration (1998). In ACIAR Proceedings of the Conference on Sandal, Bangalore, 18–19 December 1997, pp. 69–73.

  • Woodall, G. S. and Robinson, C. J., Same day plantation establishment of the ischolar_main hemiparasite sandalwood Santalum spicatum (R.Br.) A D. C: Santalaceae and its hosts. J. R. Soc., W. Aust., 2002, 85, 37–42.

  • Arun Kumar, A. N., Geeta, J. and Mohan Ram, H. Y., Sandalwood: history, uses and present status and the future. Curr. Sci., 2012, 103(12), 1408–1416.

  • Nageswara Rao, M., Soneji, J. R. and Sudarshana, P., Santalum. In Wild Crop Relatives: Genomic and Breeding Resources (ed. Kole, C.), Springer, Berlin, Germany, 2011, pp. 131–144; doi:10.1007/ 978-3-642-21250-5.

  • Radomiljac, A. M., Anathapadmanabha, H. S., Welbourn, R. M. and Satyanarayana Rao, K., The effect of sandalwood availability on the craftsman community. In Sandal and its Products. ACIAR Proceedings (84), Australian Centre for International Agricultural Research, Canberra, Australia, 1998, p. 204.

  • Heena Kausar, S., Jahan, N., Ahmed, K., Aslam, M., Ahmed, P. and Ahmed, S., Unani perspective and recent studies of sandal safed (Santalum album Linn.): a review. World J Pharm. Sci., 2014, 3, 2133–2145.

  • Sen Sarma, P. K., Sandalwood – its cultivation and utilization. In Cultivation and Utilization of Aromatic Plants (eds Atal, C. K. and Kapur, B. M.), Regional Research Laboratory, CSIR, Jammu Tawi, 1982, pp. 395–495.

  • Troup, R. S., The Silviculture of Indian Trees, Clarendon Press, Oxford, UK, 1921, vol. 3, p. 817.

  • Neil, P. E., Growing sandalwood in Nepal: potential silvicultural methods and research priorities. USDA Forest Service General Technical Report, 1990, p. 122.

  • Barber, C. A., Studies in ischolar_main parasitism – the houstorium of Santalum album L. 1. Early stages up to penetration. Mem. Dept. Agric. Ind. Bot. Ser. 1, 1903, 1, 1–26.

  • Barber, C. A., Studies in ischolar_main parasitism: the haustoria of Santalum album L. Mem Dept. Agric. India Bot. Ser. 1 Part 1, 1906, 1, 1–26.

  • Taide, Y. B., Babu, L. C. and Abraham, C. C., Influence of host species on the initial growth and development of sandal (S. album L.). Indian J. For., 1994, 174, 288–292.

  • Rocha, D., Ashokan, P. K., Santhoshkumar, A. V., Anoop, E. V. and Sureshkumar, P., Anatomy and functional status of haustoria in field-grown sandalwood tree (Santalum album L.). For. Res., 2015, 148; doi:10.4172/2168-9776.1000148.

  • Kuijt, J., The Biology of Parasitic Flowering Plants, University of California Press, Berkeley, CA, USA, 1969.

  • Barrett, D. and Fox, J. E. D., Early growth of santalum album relation to shade. Aust. J. Bot., 1994, 42, 83–93.

  • Barrett, D. and Fox, J. E. D., A protocol suitable for raising seedlings of S. album in Timor. In Sandalwood Seed Nursery and Plantation Technology (eds Gyerum, L., Fox, J. E. D. and Ehrhart, L.), Field Document. FAO, Suva, Fiji, 1995, pp. 163–164.

  • Sreenivasan Rao, Y. V., Contributions to the physiology of sandal (Santalum album L.). Influence of the host plants on the nitrogen metabolism of sandal. J. Indian Inst. Sci., 1933, 16, 164–184.

  • Iyengar, A. V. V., The relation of soil nutrients to the incidence of spike disease in sandalwood (Santalum album Linn.). Indian For., 1960, 86, 220–230.

  • Rangaswamy, C. R., Jain, S. H. and Parthasarthi, K., Soil properties of some sandalwood bearing areas. Van Vigyan, 1986, 24, 61–68.

  • Parthasarathi, K. and Rao, P. S., Studies on sandal spike: physiological significance of disturbed iron balance in the spike disease of sandal (S. album L.). Proc. Indian Acad. Sci., Sect. B, 1962, 55, 99–106.

  • Srimathi, R. A., Babu, D. R. C. and Sreenivasaya, M., Influence of host plant on the amino acid make up of Santalum album. Curr. Sci., 1961, 30(11), 417.

  • Radomiljac, A. M., McComb, J. A. and McGrath, J. F., Intermediate host influences on the ischolar_main hemi-parasite Santalum album L. biomass partitioning. For. Ecol. Manage., 1999, 113, 143–153.

  • Ehrhart, Y. and Fox, J. E. D., State of knowledge regarding cultivation of sandalwood. In Sandalwood Seed Nursery and Plantation technology (eds Gjerum L., Fox, J. E. D. and Ehrhart, Y.), Proceedings of a Regional Workshop for Pacific Island Countries, Noume’a, New Caledonia, FAO, Suva, Fiji, 1–11 August 1994, pp. 275–291.

  • Fox, J. E. D., Doronila, A. I., Barrett, D. R. and Surata, I. K., Desmanthus virgatus (L.) Willd. An efficient intermediate host for the parasitic species Santalum album L. in Timor, Indonesia. J. Sustain For., 1996, 3, 13–23; doi:10.1300/J091v03n04_02.

  • Rao, M. R., Host plants of the sandal tree. Indian For., 1911, 2, 159–207.

  • Nagaveni, H. C. and Srimathi, R. A., Studies on germination of sandal (Santalum album L.) pre-treatment of sandal seeds. Indian For., 1981, 107, 348–354.

  • Nagaveni, H. C. and Srimathi, R. A., A note on haustoria-less sandal plants. Indian For., 1985, 111, 615–618.

  • Barrett, D. and Fox, J. E. D., Santalum album: kernel composition, morphological and nutrient characteristics of pre-parasitic seedlings under various nutrient regimes. Ann. Bot., 1997, 79, 59–66; doi:10.1006/anbo.1996.0303.

  • Surata, K., Effect of host plants on growth of sandalwood (Santalum album) seedlings. Santalum, 1992, 9, 1–10.

  • Shinde, S., Ghatge, R. and Mehetre, S., Comparative studies on the growth and development of sandalwood tree in association with different hosts. Indian J. For., 1993, 16, 165–166.

  • Fox, J. E. D., Doronila, A. I. and Barrett, D. R., The selection of superior pot hosts for maximum nursery growth in Santalum album. Papers presented at the Sandalwood workshop held at Noumea, New Caledonia, Australian Centre for International Agricultural Research, CIRAD/FAO, 12 August 1994, p. 111.

  • Rai, S. N., Status and cultivation of sandalwood in India. In Proceedings of the Symposium on Sandalwood in the Pacific (eds Hamilton L. and Conrad, C. E.), USDA Forest Service General Technical Report, PSW-122, USA, 1990, pp. 66–71.

  • Nagaveni, H. C. and Vijayalakshmi, G., Growth performance of sandal (Santalum album L.) with different host species. Sandalwood Res. Newsl., 2003, 18, 1–4.

  • Venkata Rao, M. G., The influence of host plants on sandal spike disease. Indian For., 1938, 64, 656–669.

  • Ananthapadmanabha, H. S., Nagaveni, H. C. and Rai, S. N., Influence of host plants on the growth of sandal. My For., 1988, 24, 154–160.

  • Nagaveni, H. C. and Vijiyalakshmi, G., Growth performance of sandal (Santalum album L.) with different leguminous and nonleguminous host species. Sandalwood Res. Newsl., 2004, 18, 1–4.

  • Nagaveni, H. C. and Vijayalakshmi, G., Differential response in the haustorial formation and growth of sandalwood plants (Santalum album Linn.) with respect different hosts. In National Seminar on Conservation, Improvement, Cultivation and Management of Sandal, IWST Bengaluru, 12–13 December 2007, pp. 137–142.

  • Irawan, T., A study report on the effect of host diversity on growth and development of sandalwood in Kupang. In Thomas Lion (2017), Sandalwood cultivation. Int. J. Dev. Res., 2017, 7(8), 14826–14830.

  • Deepa, P. and Yusuf, A., Influence of different host associations on glutamine synthetase activities and ammonium transporter in Santalum album L. Physiol. Mol. Biol. Plants, 2016, 22(3), 331– 340.

  • Li, Y. L., Sandalwood Introduction and Research (in Chinese), Science Press, Beijing, China, 2003, p. 3.

  • Matthies, D., Parasite–host interaction in Castillega and Orthocarpus. Can. J. Bot., 1997, 751, 1252–1260.

  • Barrett, D. J., Hatton, T. J., Ash, J. E. and Ball, M. C., Transpiration by trees from contrasting forest types. Aust. J. Bot., 1996, 44, 249–263.

  • Brand, J. E., Crombie, D. S. and Mitchell, M. D., Establishment and growth of sandalwood (Santalum spicatum) in south-western Australia: the influence of host species. Aust. For., 2000, 63, 60– 65; doi:10.1080/00049158.2000.10674814.

  • Brand, J. E., Ryan, P. C. and Williams, M. R., Establishment and growth of sandalwood (Santalum spicatum) in south–western Australia: the Northampton pilot trial. Aust. For., 1999, 62, 33–37.

  • Barrett, D. R., Wijesuriya, S. R. and Fox, J. E. D., Observations on foliar nutrient content of sandalwood (Santalum spicatum R.Br. D.C.). Mulga Res. Centre J., 1985, 8, 81–91.

  • Barrett, D. R., Santalum album (Indian sandalwood) literature survey. Mulga Res. Centre, Curtin University, Perth, Australia, 1988, p. 5.

  • Sen-Sarma, P. K., Some aspects of ecology of sandal spike disease. In Proceedings of All India Sandal Seminar, Karnataka Forest Department, Bangalore, 1977, pp. 163–166.

  • Sreenivasan, V. V., Shivaramakrishnana, V. R., Rangaswamy, C. R., Anathapadmanabha, H. S. and Shankaranarayana, K. H., Sandal, ICFRE, Dehra Dun, 1992, p. 233.

  • McKinell, F. H., Status of management and silviculture research on sandalwood in Western Australia and Indonesia. In Proceedings of the Symposium on Sandalwood in the Pacific (eds Hamilton, L. and Conrad, C. E.), United States Department of

  • Agriculture, General Technical Report, PSW-122, Hawaii, 1990, pp. 19–25.

  • Viswanath, S., Dhanya, B., Purushothaman, S. and Rathore, T. S., Financial viability of sandal (Santalum album) based agroforestry practices in Southern India. Indian J. Agrofor., 2010, 12(2), 14– 22.

  • Jonathan, E. B., Len, J. N. and Ian, C. D., Estimated heartwood wiehgts and oil concentrations within 16 year old Indian sandalwood (Santalum album) trees planted near Kununurra, Wesstern Australia. Aust. For., 2012, 75(4), 225–232.

  • Kelly, C. K., Venable, D. L. and Zimmerer, K., Host specialization in Cuscuta costaricensis: an assessment of host use relative to host availability. Oikos, 1988, 53, 315–320.

  • Tennakoon, K. U., Ekanayake, S. P. and Etampawala, L., An introduction and current status of sandalwood research in Sri Lanka. Int. Sandalwood Res. Newsl., 2000, 10, 1–4.

  • Wawo, A. H., Cendana, Deregulation and Strategy, Pengembangannya, Bogor (2002). In Thomas Lion (2017) Sandalwood cultivation. Int. J. Dev. Res., 2017, 7(8), 14826–14830.

  • Cameron, D. D., Hwangbo, J. K., Keith, A., Geniez, J. M., Kraushaar, D., Rowntree, J. and Seel, W. E., Interactions between the hemiparasitic angiosperm Rhinanthus minor and its hosts: from the cell to the ecosystem. Folia Geobot., 2005, 40, 217– 229.

  • Varghese, S., Parasitic interference of sandal (Santalum album Linn.) on common agricultural crops from the homesteads. M.Sc. thesis, Kerala Agricultural University, Thrissur, 1997.

  • Vijayakumar, H., Hiremath, J. and Ashokan, P. K., Influence of soil moisture regimes and stage of host introduction on seedling growth of sandal provenances. In National Seminar on Conservation, Improvement, Cultivation and Management of Sandal, IWST Bengaluru, 12–13 December 2007, pp. 59–66.

  • Dhaniklal, G., Influence of host plants on soil moisture stress on the water relations in sandal. M.Sc. thesis, Kerala Agricultural University, Thrissur, 2006.

  • Ashokan, P. K. and Krishnambika, N., Growing sandal in home garden and other agroforestry systems – potentials and problems. In Proceedings of a National Seminar, IWST, Bangalore, 12–13 December 2007, pp. 143–152.

  • Rocha, D., Ashokan, P. K., Santhoshkumar, A. V., Anoop, E. V. and Sureshkumar, P., Influence of host plant on the physiological attributes of field-grown sandal tree (Santalum album). J. Trop. For. Sci., 2014, 26, 166–172.

  • Pate, J. S., Kuo, J. and Davidson, N. J., Growth, resource allocation and haustorial biology of the ischolar_main hemiparasite Olax phyllanthi (Olacaceae), with special reference to the haustorial interface. Ann. Bot., 1990, 65, 425–436.

  • Babita, M., Sandeep, C., Sandhya, M. C. and Viswanath, S., Sandalwood farming in India: problems and prospectus. Indian J. Trop. Biodivers., 2018, 26(1), 1–12.

  • Ananthapadmanabha, H. S., Indian sandalwood market trend and production. In International Seminar on Sandalwood: Current Trends and Future Prospects (ed. Ramakantha, V.), Institute of Wood Science and Technology, Indian Council of Forestry Research and Education, Bangalore, 2014, p. 74.

  • Viswanath, S., Ahmed, S. M., Madhu, K. S., Kumar, N. P., Sowmya, C., Arade, S. C. and Adhikari, M., Performance of sandalwood based agroforestry models with horticulture crops as secondary hosts in Karnataka. Abstracts of International Seminar on Sandalwood: Current Trends and Future Prospects, IWST, Bangalore, 26–28 February 2014, p. 24.

  • Singh, B., Rathore, T. S. and Singh, G., The effect of long term hosts species on Santalum album L. growth under agroforestry in semi-arid condition of North Gujarat, India. In Abstracts of International Seminar on Sandalwood: Current Trends and Future Prospects, IWST, Bangalore, 26–28 February 2014, p. 26.

  • Hibberd, J. M., Quick, W. P., Press, M. C. and Scholes, J. D., The influence of the parasitic angiosperm Striga gesnerioides on the growth and photosynthesis of its host, Vigna unguiculata. J. Exp. Bot., 1996, 47, 507–512.

  • Fisher, J. P., Phoenix, G. K., Childs, D. Z., Press, M. C., Smith, S. W., Pilkington, M. G. and Cameron, D. D., Parasitic plant litter input: a novel indirect mechanism influencing plant community structure. New Phytol., 2013, 198, 222–231.

  • Press, M. C., Scholes, J. D. and Watling, J. R., Parasitic plants: physiological and ecological interactions with their hosts. In Physiological Plant Ecology (eds Press, M. C., Scholes, J. D. and Barker, M. G.), Blackwell, Science, Oxford, UK, 1999, pp. 175– 197.

  • Ananthapadmanabha, H. S., Rangaswamy, C. R., Sarma, C. R., Nagaveni, H. C., Jain, S. H., Venkatesan, K. R. and Krishnappa, H. P., Host requirement of sandal (Santalum album L.). Indian For., 1984, 110(3), 264–268.

  • Barbour, L., Analysis of plant–host relationships in tropical sandalwood (Santalum album). Rural Indust. Res. Develop. Corp., Kununurra, Western Australia, 2008, 8(138), pp. 1–25.

  • Doddabasawa, Chittapur, B. M. and Lokesh, R., Parasitism ecology of sandalwood (Santalum album L.) for commercial production in the semi-arid tropics. Curr. Sci., 2020, 119(4), 699– 703.

  • Teixeira da Silva, J. A., Kher, M. M., Soner, D., Page, T., Zhang, X., Nataraj, M. and Ma, G., Sandalwood: basic biology, tissue culture and genetic transformation. Planta, 2016, 243, 847–887.

  • Brand, J. E. and Jones, P., Growing sandalwood (Santalum spicatum) on farmland in Western Australia. Forest Products Commission, Western Australia, Perth, Australia, 2001, pp. 1–25.

  • Annapurna, D., Rathore, T. S. and Joshi, G., Modern nursery practices in the production of quality seedlings of Indian sandalwood (Santalum album L.) – stage of host requirement and screening of primary host species. J. Sustain For., 2006, 22, 33–55; doi:10.1300/J091v22n03_03.

  • Davit, M. W., Determinants of parasitic plant distribution: the role of host quality. Botany, 2009, 87(1), 16–21.

  • Rai, S. N. and Sarma, C. R., Study of diameter growth in sandal. J. Trop. For., 1986, 2, 202–206.

  • Iyenger, A. V. V., The physiology of ischolar_main parasitism in sandal (Santalum album L.). Indian For., 1965, 91, 246–256.

  • McComb, J. A., Clonal Santalum album growth, oil content and composition on different hosts and at different locations. J. Roy. Soc. W. Aust., 2009, 92, 15–25.

  • Rocha, D., Ashokan, P. K., Santhoshkumar, A. V., Anoop, E. V. and Sureshkumar, P., Anatomy and functional status of haustoria in field grown sandalwood tree (Santalum album L.). Curr. Sci., 2017, 113(1), 130–133.

  • Parthasarathi, K., Gupta, S. K. and Rao, P. S., Differential response in the cation exchange capacity of the host plants on parasitization on sandal (Santalum album). Curr. Sci., 1974, 43, 17–20.

  • Ma, G. H., He, Y. M., Zhang, J. F. and Chen, F. L., Studies on semi-parasitic sandalwood seedlings. J. Trop. Subtrop. Bot., 2005, 13, 233–238.

  • Lu, J. K., The parasitism between hemiparasite Santalum album and its hosts. Ph.D. thesis, Chinese Academy of Forestry, Beijing, China, 2011, p. 37.

  • Lu, J. K., Kang, L. H., Sprent, J. I., Xu, D. P. and He, X. H., Two way transfer of nitrogen between Dalbergia odorifera and its hemiparasite Santalum album is enhanced when the host is effectively nodulated and fixing nitrogen. Tree Physiol., 2013, 33, 464– 474.

  • Lu, J., Xu, D., Kang, L. and He, X., Host-species-dependent physiological characteristics of hemiparasite Santalum album in association with N2-fixing and non-N2-fixing hosts native to southern China. Tree Physiol., 2014, 34, 1006–1017.

  • Zhang, X. H., Teixeira da Silva, J. A., Duan, J., Deng, R., Xu, X. and Ma, G. H., Endogenous hormone levels and anatomical characters of haustoria in Santalum album L. seedlings before and after attachment to the host. J. Plant Physiol., 2012, 169, 859– 866; doi:10.1016/j.jplph.2012.02.010.

  • Chen, R., Zhang, X. H. and Ma, G. H., Studies on parasitic relationship between Santalum album L and leguminous plants. J. Trop. Subtrop. Bot., 2014, 22, 53–60.

  • Ouyang, Y., Zhang, X. H., Chen, Y. L., Teixeira da Silva, J. A. and Ma, G. H., Growth, photosynthesis and haustorial development of semiparasitic Santalum album L. penetrating into ischolar_mains of three hosts: a comparative study. Trees – Struct. Funct., 2015, 1–13; doi:10.1007/s00468-015-1303-3.

  • Reddy, G. R. S., Honnuri, M. B. and Lokeshwara Rao, M., Evaluation of performance of sandal (Santalum album L.) germplasm under agroforestry conditions at Hyderabad. In National Seminar on Conservation, Improvement, Cultivation and Management of Sandal (Santalum album L.), IWAST, Bengaluru, 12–13 December 2007, pp. 107–105.


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  • Sandalwood Plantations – Points to Ponder

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Authors

Doddabasawa
Division of Farm Forestry, College of Agriculture, Bheemarayanagudi, Yadgir 585 287, India
B. M. Chittapur
Directorate of Extension, University of Agricultural Sciences, Raichur 584 104, India

Abstract


East Indian sandalwood (Santalum album L.) is one of the precious woods known for sweet fragrant aroma and commercial value in both national and international markets. Besides, its wide adaptability to varied climate, hosts and edaphic conditions have attracted farmers and corporates for commercial venture. Sandalwood being a hemi-ischolar_main parasite, the successful establishment of sandalwood plantation depends on the understanding of parasitism ecology, especially the relations between host and parasite, their ratio and other silviculture techniques. Hence, the present article aims to understand these issues for commercial upscaling. Sandalwood relies largely on host plants for mineral nutrients and water through haustorial connections which act as a physiological and physical bridge between the parasite and the host. It parasitizes over a vast array of plants from grasses to trees, whereas leguminous associations are superior. Deepischolar_mained and slow-growing perennial hosts help in sustained growth. For better yield of heartwood and oil sandalwood should be grown over 15 years, whereas optimum rotation age would be 25–30 years. Sandalwood can be planted either through direct dibbling of seeds near the hosts in the same pit or adjacent to the hosts. Better establishment occurs when sandalwood is grown with potted host followed by secondary hosts in the field. Association of sandalwood and host would be strong when the distance is around 2.5–3 m with host to parasitic ratio of 2 : 1 or 3 : 1. However, questions on parasitism ecology still remain.

Keywords


Ecology, Haustoria, Hosts, Parasite, Sandalwood.

References





DOI: https://doi.org/10.18520/cs%2Fv120%2Fi7%2F1184-1193